Boston Properties’ executive bonuses rose as the commercial developer racked up 7.2 million square feet of leasing activity in 2018.

Executive Vice President Bryan Koop, who heads the company’s Greater Boston operations, saw his bonus rise 21 percent to $1.55 million while his salary was unchanged at $400,000, Boston Properties reported in its annual proxy statement. Koop received $1.3 million in stock options plus $34,380 in other compensation, bringing his total compensation up to $3.2 million.

The company secured anchor tenants for four office developments totaling more than 2.1 million square feet including Verizon Communications’ 440,000 square foot lease at The Hub on Causeway and 638,000 square feet by Google at the Kendall Center development in Cambridge.

Online security firm Mimecast relocated from Watertown to 191 Spring St. in Lexington following a repositioning of the 200,000-square-foot office building, replacing previous tenant Wolverine Worldwide. The Proto apartment tower in Kendall Square was 56 percent leased as of Dec. 31, Boston Properties said. And its 1.4-million-square-foot Salesforce Tower in San Francisco, which opened in late 2018, is 100 percent leased.

Boston Properties said the 7.2 million-square-foot leasing pipeline, the second-highest in company history, represents nearly 15 percent of its portfolio. The company owns property in the Boston, New York, San Francisco, Los Angeles and Washington, D.C. metro areas.

CEO Owen Thomas’ total compensation rose to $11.7 million, up from $10 million in 2017. Thomas’ compensation represents a 107-to-1 ratio compared to the company’s median pay for all 747 employees, which is $109,173.